| W | Width of the alignment |
| N | Number of sequences in the alignment |
| cki | Count of nucleotide i in the kth column of the alignment |
| pi | Background probability of nucleotide i |
| αi | ith pseudocount |
| A | Sum of pseudocounts (pseudocount weight) |
| Γ | Gamma function |
... Aren't you glad you asked?
For an explanation see our hopefully soon to be published GLAM paper.